• Wascher, C.A.F., Allen, K. & Szipl, G. 2021. Learning and motor inhibitory control in crows and domestic chickens. Royal Society Open Science 8: 210504210504
  • Wascher, C.A.F. 2021. Heart rate as a measure of emotional arousal in evolutionary biology. Philosophical Transactions of the Royal Society B 376: 20200479. (link to open access pre-print)
  • Khan, N., & Wascher, C.A.F. 2021. Considering generalizability: A lesson from auditory enrichment research on zoo animals.  Animal Behavior and Cognition, 8(2), 251-262.  (link to pdf)
  • Wascher C.A.F. 2021. Association between social factors and gastrointestinal parasite product excretion in a group of non-cooperatively breeding carrion crows. Behavioral Ecology and Sociobiology, 75, 30. (link to open access pre-print)
  • Dufour V., Broihanne M.-H. & Wascher C.A.F. 2020. Corvids avoid odd evaluation by following simple rules in a risky exchange task. Ethology, 126(2), 153-164. (link to pdf)
  • Wascher C.A.F., Feider B., Bugnyar T. & Dufour V. 2020. Crows and common ravens do not reciprocally exchange tokens with a conspecific to gain food rewards. Ethology, 126(2), 278-287. (link to pdf)
  • Wascher C.A.F., Canestrari D. & Baglione V. 2019. Affiliative social relationships and coccidian oocyst excretion in a cooperatively breeding bird species. Animal Behaviour, 158, 121-130. (link to open access pre-print)
  • Halsey L.G., Green J.A., Twiss S.D., Arnold W., Burthe S.J., Butler P.J., Cooke S.J., Grémillet D., Ruf T., Hicks O., Minta K.J., Prystay T.S., Wascher C.A.F., & Careau V. 2019. Flexibility, variability and constraint in energy management patterns across vertebrate taxa revealed by long-term heart rate measurements. Functional Ecology, 33 (2). 260-272. (send e-mail to request pdf)
  • Frigerio D., Hemetsberger F. & Wascher C.A.F. 2019. How having a partner and a family changes the stress levels of graylag geese. Frontiers for Young Minds, 7, 68. doi: 10.3389/frym.2019.00068. (link to pdf)
  • Safryghin A., Hebesberger D.V. & Wascher C.A.F. 2019. Testing for behavioral and physiological responses of domestic horses (Equus caballus) across different contexts – consistency over time and effects of context. Frontiers in Comparative Psychology 10, 849, (link to pdf)
  • Miller R., Boeckle M., Jelbert S.A., Frohnwieser A., Wascher C.A.F. & Clayton N.S. 2019. Self-control in crows, parrots and non-human primates. WIREs Cognitive Science, e1504. (link to pdf)
  • Szipl G., Loth A., Wascher C.A.F., Hemetsberger J., Kotrschal K. & Frigerio D. 2019. Parental behaviour and family proximity as key to gosling survival in Greylag geese (Anser anser). Journal of Ornithology, 160, 473–483. (link to open access pre-print)
  • Broihanne M.H., Romain A., Call J., Thierry B., Wascher C.A.F., De Marco A., Verrier D. & Dufour V. Monkeys (Sapajus apella and Macaca tonkeana) and great apes (Gorilla gorilla, Pongo pygmaeus, Pan paniscus and Pan troglodytes) play for the highest bid. Journal of Comparative Psychology, in press. (link to open access pre-print)
  • Wascher C.A.F. †, Kulahci, I.G. †, Langley E. & Shaw R. 2018. How does cognitive performance shape patterns of social relationships? Philosophical Transactions of the Royal Society B 373, 20170293.  †These authors contributed equally to this study. (link to open access pre-print)
  • Kirk J. & Wascher C.A.F. 2018. Temporal modification of social interactions in response to changing group demographics and offspring maturation in African lions (Panthera leo). Behavioural Processes, 157, 519-527. (send e-mail to request pdf)
  •   Puehringer-Sturmayr V., Wascher C.A.F., Loretto M.-C., Palme R., Stoewe M., Kotrschal K. & Frigerio D. 2018 Seasonal differences of corticosterone metabolite concentrations and parasite burden in northern bald ibis (Geronticus eremita): The role of affiliative interactions. PLoS ONE, 13(1), e0191441. (link to pdf)
  • Wascher C.A.F., Kotrschal K. & Walter A. 2018. Free-living Greylag geese adjust their heart rates and body core temperatures to season and reproductive context. Scientific Reports, 8, 2142. (link to pdf)
  • Wascher C.A.F., Hemetsberger J., Kotrschal K. & Frigerio D. 2017. Leucocyte profiles and social context in juvenile greylag geese (Anser anser). Avian Biology Research, 10 (4), 246–252. (link to pdf)
  • Ludwig S.C., Kapetanopoulos K., Kotrschal K. & Wascher C.A.F. 2017. Mate separation induces immunosuppression and increases parasite load in free-living female Greylag geese. Behavioural Processes, 138, 134–141. (link to pdf)
  • Frigerio D., Ludwig S.C., Hemetsberger J., Kotrschal K. & Wascher C.A.F. 2017. Social and environmental factors modulate leucocyte profiles in free-living Greylag geese (Anser anser). PeerJ 5, e2792; DOI 10.7717/peerj.2792. (link to pdf)
  • Frigerio D., Cibulski L., Ludwig S.C., Camderrich I., Kotrschal K. & Wascher C.A.F. 2016. Excretion patterns of coccidian oocysts and nematode eggs during the reproductive season in Northern Bald Ibis (Geronticus eremita) Journal of Ornithology, 157, 839-851. (link to pdf)
  • Wascher C.A.F., Hillemann F., Canestrari D., & Baglione V. 2015. Carrion crows learn to discriminate between calls of reliable and unreliable conspecifics. Animal Cognition, 18, 1181-1185. (link to pdf)
  • Szipl G., Boeckle M., Spreafico M., Wascher C.A.F. & Thomas Bugnyar T. 2015. With whom to dine? Common ravens are attracted by yells of affiliates and consider signallers rank. Animal Behaviour 99, 33-42. (link to pdf)
  • Wascher C.A.F. 2015. Individual performance in socio-cognitive tasks predicts social performance in carrion crows. Behaviour 152, 615-634. (link to pdf)
  • Wascher C.A.F., Heiss, R., Baglione V. & Canestrari D. 2015. Behavioural responses to olfactory cues in social contexts in carrion crows (Corvus corone corone). Behavioural Processes 111, 1-5. (send e-mail to request pdf)
  • Dorn S., Wascher C.A.F. & Kotrschal, K. 2014. Ambient temperature and air pressure modulate hormones and behavior in Greylag geese (Anser anser) and Northern bald ibis (Geronticus eremita). Behavioural Processes 108, 27-35. (link to pdf)
  • Cibulski L., Wascher C.A.F., Weiß B.M. & Kotrschal, K. 2014. Familiarity with the experimenter influences the performance of Common ravens (Corvus corax) and Carrion crows (Corvus corone corone) in cognitive tasks. Behavioural Processes 103, 129-137. (link to pdf)
  • Hillemann, F., Bugnyar, T., Kotrschal, K. & Wascher, C.A.F. 2014. Waiting for better, not for more: corvids respond to quality in two delay maintenance tasks. Animal Behaviour 90, 1-10. (link to pdf)     
  • Wascher C.A.F., Valdez J.W., Nuñez C., Baglione V. & Canestrari D. 2014. Social factors modulating attention patterns in carrion crows (Corvus corone corone). Behaviour 151 (5), 555-572. (link to pdf)     
  • Teschke I., Wascher C.A.F., Scriba M., von Bayern A., Huml V., Siemers B. & Tebbich S. 2013. Did tool-use evolve with enhanced physical cognitive abilities? Philosophical Transactions of the Royal Society B 368, 20120418. (link to pdf)  
  • Wascher C.A.F. & Bugnyar T. 2013. Awareness to inequity and working effort in carrion crows (Corvus corone corone) and common ravens (Corvus corax). PLoS ONE 8(2), e56885. (link to pdf)  
  • Wascher C.A.F., Bauer A.C., Holtmann A.R. & Kotrschal K. 2012. Environmental and social factors affecting excretion of intestinal parasite eggs and oocysts in a flock of free-living greylag geese (Anser anser). Behavioral Ecology 23 (6), 1276-1283. (link to pdf)  
  • Wascher C.A.F., Dufour V. & Bugnyar T. 2012. Carrion crows cannot overcome impulsive choice in a quantitative exchange task. Frontiers in Comparative Psychology 3, 118. (link to pdf)  
  •  Wascher C.A.F., Szipl G., Boeckle M. & Wilkinson A. 2012. You sound familiar- Carrion crows can differentiate between the calls of known and unknown heterospecifics. Animal Cognition 15, 1015-1019. (link to pdf)  
  • Wascher C.A.F., Weiß B.M., Arnold W. & Kotrschal K. 2012. Physiological implications of pair-bond status in greylag geese. Biology Letters 8, 347-350. (link to pdf)  
  • Dufour V., Wascher C.A.F., Braun A., Miller R. & Bugnyar T. 2012. Corvids can decide if a future exchange is worth waiting for. Biology Letters 8, 201-204. (link to pdf)  
  • Wascher C.A.F., Scheiber I .B.R., Braun A. & Kotrschal K. 2011. Heart rate responses to induced challenge situations in greylag geese (Anser anser). Journal of Comparative Psychology 125 (1), 116-119. (link to pdf)  
  • Wascher C.A.F., Fraser O.N. & Kotrschal K. 2010. Heart rate during conflicts predicts post-conflict stress-related behavior in greylag geese. PLoS ONE 5 (12), e15751. (link to pdf)  
  • Kralj-Fiser S., Scheiber I.B.R., Kotrschal K., Weiß B.M. & Wascher C.A.F. 2010. Glucocorticoids enhance and suppress heart rate and behaviour in time dependent manner in greylag geese (Anser anser). Physiology & Behavior 100, 394-400. (link to pdf)  
  • Wascher C.A.F., Scheiber I .B.R., Weiß B.M. & Kotrschal K. 2009. Heart rate responses to agonistic encounters in greylag geese (Anser anser). Animal Behaviour 77, 955-961. (link to pdf
  • Scheiber I.B.R., Weiß B.M., Hirschenhauser K., Wascher C.A.F., Nedelcu I.T. & Kotrschal K. 2008. Does 'relationship intelligence' make big brains in birds? The Open Biology Journal 1, 6-8. (link to pdf
  • Wascher C.A.F., Arnold W. & Kotrschal K. 2008. Heart rate modulation by social contexts in Greylag geese (Anser anser). Journal of Comparative Psychology 122 (1), 100-107. (link to pdf
  • Wascher C.A.F., Scheiber I.B.R. & Kotrschal K. 2008. Heart rate modulation in bystanding geese watching social and non-social events. Proceedings of the Royal Society London, Series B 275, 1653-1659. (link to pdf

Book chapters

·       Wascher C.A.F. 2018. Corvids. In: Encyclopedia of Animal Cognition and Behavior, eds. Vonk J. and Shackelford, T.K. Springer. Doi:

·       Wascher C.A.F. & Kotrschal K. 2013. The costs of sociality measured through heart rate modulation. In: The Social Life of Greylag Geese. Patterns, Mechanisms and Evolutionary Function in an Avian Model System, eds. I.B.R. Scheiber, B. Weiß, J. Hemetsberger and K. Kotrschal. Cambridge: Cambridge University Press: 142-155. ISBN: 9780521822701.

                            Published abstracts

·       Ludwig S.C., Kapetanopoulos K., Wascher C.A.F. & Kotrschal K. 2012. Effects of mate separation on stress hormones, parasite load and immune parameters in free-living greylag geese. Wiener Tierärztliche Monatsschrift 99 (Suppl 1), 61. ISSN 0043-535x

·       Spreafico M, Szipl G., Kotrschal K. & Wascher C.A.F. 2012. Physiological and behavioural response in carrion crows (Corvus corone corone) after relocation to a new environment. Wiener Tierärztliche Monatsschrift 99 (Suppl 1), 61. ISSN 0043-535x.

·       Szipl G., Wascher C.A.F., Spreafico M., Barth E.K., Kotrschal K. & Bugnyar T. 2012. Vocal response to stressful situations in carrion crows (Corvus corone corone). Wiener Tierärztliche Monatsschrift 99 (Suppl 1), 62. ISSN 0043-535x.

·       Scheiber I.B.R., Wascher C.A.F. & Kotrschal K. 2006. Biparental care in Greylag Geese: Heart rate as a measure of parental contribution. Journal of Ornithology 147 (Suppl 1), 245. ISSN: 2193-7192.

·       Wascher C.A.F. 2010. Heart rate in greylag geese (Anser anser): implications for the study of social systems. Etho-News: 64, 9.

·       Wascher C.A.F, Kralj-Fiser S., Scheiber I.B.R. & Kotrschal K. 2006. Heart rate and coping styles in Greylag Geese. Journal of Ornithology 147 (Suppl 1), 269. ISSN: 2193-7192.